1. walking or standing upright on two legs
Usage: biology; anatomy

Examples

• Humans are orthograde primates that walk upright on two legs.
• The orthograde posture of early hominids was a key evolutionary development.
• Unlike quadrupedal mammals, orthograde species have adapted spines for vertical support.
• The transition from pronograde to orthograde locomotion marked human evolution.
• Orthograde walking requires significant changes in skeletal structure.
• Birds are orthograde when standing but not when walking.